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In the winter of 1152, Castle Keep was on the brink of survival due to a lack of food and medicine. Several groups of guards have been sent as envoys to other mouse cities to build cooperation and get supplies.

Saxon, Kenzie, Liam, and Sadie, along with Selanav, the legendary Black Axe, travel through the snowy lands and race against time to deliver the Spruce Elixir to the harbour. Not every guardian will be able to survive this winter.